How has evolution molded human biological evolution and cultural diversity? What are the broader implications of our evolutionary history? This class seeks answers to these enduring questions. Divided into four parts, we begin by examining Darwin's ideas about natural selection and the challenges his ideas have faced over the past 150 years. The second section examines how evolution has shaped the anatomy and behavior of our closest living relatives, non-human primates. The third part of the class is devoted to investigating the human fossil record and tracing the physical and behavioral evolution of our species over the past five million years. In the fourth section, we focus on the breadth of human variation and explore the broader social implications of our evolved behavior.